About Suncus etruscus (Savi, 1822)
The Etruscan shrew (Suncus etruscus, first described by Savi in 1822) has a slender, non-truncated body. Its body length (excluding the tail) ranges from 3 to 5.2 cm (1.2 to 2.0 in), while the tail adds an additional 2.4 to 3.2 cm (0.94 to 1.26 in). Body mass falls between 1.3 g (0.046 oz) and 2.5 g (0.088 oz), with an average mass of around 1.8 g (0.063 oz). The related greater white-toothed shrew is roughly twice as long and weighs four to five times more than the Etruscan shrew. The Etruscan shrew has a relatively large head, a long mobile proboscis, relatively small hind limbs, and relatively large, prominent ears. It has an extremely fast heart rate that can reach up to 1511 beats per minute (25 beats per second), and a relatively large heart muscle mass that makes up 1.2% of its total body weight. Its fur is pale brown on the back and sides, and light gray on the stomach. Fur becomes denser and thicker from fall through the winter. Etruscan shrews usually have 30 teeth, but the 4th upper intermediate tooth is very small and rudimentary, and is absent in some individuals. A dense array of short whiskers grows near the mouth, which the shrew actively uses to search for prey, particularly at night. There is no sexual dimorphism in body features between male and female Etruscan shrews. This species occupies a range that spans 10° to 40°N latitude across Eurasia. In Southern Europe, it has been confirmed in Albania, Bosnia and Herzegovina, Bulgaria, Croatia, Cyprus, France, North Macedonia, Malta, Montenegro, Greece, Italy, Portugal, Slovenia, Spain, and Turkey. Unconfirmed reports of its presence exist for Andorra, Gibraltar and Monaco. It has been introduced by humans to European islands including Mallorca and the Canary Islands. The Etruscan shrew also occurs in North Africa (Algeria, Egypt, Libya, Morocco, Tunisia) and around the Arabian Peninsula (Bahrain, Israel, Jordan, Lebanon, Oman, Syria, and Yemen, including Socotra). In Asia, it has been observed in Afghanistan, Azerbaijan, Bhutan, China (only in Gengma County), Myanmar (Burma), Georgia, India, Iran, Iraq, Kazakhstan, Laos, Malaysia (the Malaysian portion of Borneo island), Nepal, Pakistan, Philippines, Sri Lanka, Tajikistan, Thailand, Turkey, Turkmenistan and Vietnam. Unconfirmed reports note possible presence in West and East Africa (Guinea, Nigeria, Ethiopia), and in Armenia, Brunei, Indonesia, Kuwait and Uzbekistan. Overall, the species is widespread and not considered threatened, but its population density is generally lower than that of other shrews that live in the same areas. It is rare in some regions, most notably Azerbaijan, Georgia, Jordan and Kazakhstan, where it is included in the national Red Book. The Etruscan shrew prefers warm, damp habitats with shrub cover, which it uses to hide from predators. It most often inhabits areas where open terrain such as grasslands and scrub meet deciduous forests. It can be found at sea level, but is usually restricted to the foothills and lower elevation belts of mountain ranges, though it has been recorded at up to 3,000 m (9,800 ft) above sea level. It colonizes riparian thickets along the banks of lakes and rivers, as well as human-cultivated areas including abandoned gardens, orchards, vineyards, olive groves, and field edges. However, it avoids intensively cultivated areas, dense forests, and sand dunes. It is poorly adapted to digging its own burrows, so it builds nests in various natural shelters, crevices, and uninhabited burrows dug by other animals. Etruscan shrews frequently frequent rocky areas, boulders, stone walls, and ruins, darting quickly in and out between these structures.
